Two new species, the new members of the Qilianshan Fauna, referred to a new family Paranamurotypidae and Paralogidae are described from the Tupo Formation (Namurian B-C) of the Upper Carboniferous of China. Formerly Shenzhousia qilianshanensis Zhang et Hong, 2006 in family Meganeuridae was discovered from the same formation.
Odonatoptera Martynov, 1932
Paranamurotypidae fam. nov.
C basally curved; ScP extremely long; RP branched after the level of midwing; long fusion of MP with Cu; cubito-anal area narrow and long, only with 1-2 rows of cells.
The new family Paranamurotypidae can be differed easily form Erasipteridae Carpenter, 1939 and Campylopteridae Handlirsch, 1906 in ScP long, MP 'S'-shape waved, and A1 zigzagged distally. It has close affinity with the family Namurotypidae Bechly, 1996, and differs from the latter in cubito-anal area long and narrow, long fusion of MP with Cu, and distal A1 zigzagged.
Paranamurotypus xiaheyanensis gen. et sp. nov.
A forewing (part and counterpart) labeled with GMC-07QNX-003a, b, preserved 47.0 mm long, 9.5 mm wide, merely missing the tip of wing; ScA short; ScP extremely long, reaching nearly apex of wing; cross-veins sparse between C and ScP, and between ScP and RA; RP+MA separating from RA 1.5 mm from wing base, with a long arculus; RP separating from MA 14.0 mm from wing base; RP branching into RP1+2 and RP3+4 after the midwing, IR2 well defined; MA simple, only one row of cells between MA and RP3+4; MP basally separating from Cu and then fused about distance of 6.0 mm; CuA and CuP separating from MP before the curve of MP and merging into AA; MP 'S'-shape waved; cubito-anal area narrow and long, only with 1-2 rows of cells; the distal of A1 zigzagged, AP inclining to margin 40°with AA.
Paralogidae Handlirsch, 1906
Oligotypus Carpenter, 1931
Oligotypus shangheyanensis sp. nov.
Basal part of fore wing (part and counterpart) labeled with GMC-07QNS-001a, b, preserved 50.0 mm long, 20.0 mm wide, about 1/3 length of wing; precostal area 16.0 mm long; common stem of RP+MA (arculus) long; basal anal area with nine long cells longitudinally, and followed suddenly by three rows cells; AP long.
This specimen only preserved a fragment and has no enough differences from the Genus Oligotypus. So the new species referred to Oligotypus tentatively. It differs from Oligotypus tillyardi Carpenter, 1931 in the characters as stated above.
Up to now, three species of Odonatoptera have been described in the Qilianshan Fauna from the Tupo Formation of the Upper Carboniferous of China. They show a high species diversity in the group Odonatoptera. Besides the Odonatoptera, the Archaeorthoptera and the Grylloblattodea (Zhang, in prep.) also show high species diversity. It can be inferred tentatively that the Qilianshan Fauna was in a prior evolutionary stage to the Euramerican Fauna during the Carboniferous.
Supported by Programme of Excellent Young Scientists of the Ministry of Land and Resources, China.